Colonial rugose corals are common in western cratonal North America and in some of the allochthonous terranes, now amalgamated against its western margin. Throughout the Late Paleozoic, the coral faunas in these two different settings were significantly different. Comparisons of these faunas suggest that during the Mississippian the Alexander terrane probably was southwest of Arctic Alaska and the Stikine terrane probably lay west of the southern part of the North American craton. The Cache Creek terrane lay far out in the Paleopacific Ocean. The Pennsylvanian faunas suggest that the Quesnellia and Eastern Klamath terranes were situated southwest of Arctic Alaska and the Alexander terrane was somewhat farther southwest and farther from cratonal North America. The Stikine terrane continued to be positioned west of the southern part of the North American craton. During the Early Permian, terranes with a cratonal faunal aspect may have lain 2000–3000 km west of cratonal North America and latitudinally generally southwest of their present positions. In the Middle Permian these terranes were carried southward relative to the North American craton. Simultaneously the Tethyan Realm expanded eastward.